"Shinkalion" was developed by the Shinkansen Ultra Evolution Institute to protect the peace and safety of Japan from mysterious giant monstrous beings. The Ultra Evolution Institute has been developing "Shinkalion Z," which is a new type of robots, as well as "Zailiner," new armor upgrading trains that power "Shinkalion Z" up by transforming from conventional trains, to prepare for an invasion by new enemies. Children who became operators for "Shinkalion Z" with high matching rates join forces with members of the Institute, and fight back against the giant monstrous beings that emerged again! Emergence of unknown enemies... And two boys meeting each other... A further evolution of Shinkalion begins!!!